Purdue University - Extension - Forestry and Natural Resources
Seeding rates for native warm-season grass and forb mixtures (NWSG) have changed drastically over time. In the past, native grasses were planted without forbs at rates exceeding 10 lbs/ac. This may be ideal from a forage production standpoint, but this created dense stands of native grass with little to no forb component and lacked benefits to most wildlife.
Mixtures have shifted from heavy planting rates of tallgrass species with few forbs to reduced rates of mid-stature grasses with an abundance of forbs. Recommended seeding rates of some current mixtures may be lower than what no-till drills are capable of planting. In this case, fillers may be needed to increase the bulk weight of the seed to allow the equipment to plant at the correct rate.
Seed mixtures are also more commonly being established by broadcasting seed during late winter (frost seeding) using cyclone fertilizer spreaders. Broadcasting native warm-season grass and forb seed usually requires the use of a carrier to ensure the mixture flows correctly through the spreader and the seed is distributed evenly across the field.
Using fillers when no-till drilling native warm-season grass and forb mixtures
Planting native grass and forb mixtures with a no-till drill is the most common establishment method for NWSG plantings. It may be difficult to achieve the correct seeding rate with a no-till drill because of the combination of reduced bulk weight of dechaffed seed and reduced seeding rates of common mixtures. Fillers can be used to increase the bulk weight of native grass and forb seed if a drill cannot achieve the recommended seeding rate. Traditionally, native grass and forbs have been planted separately using the grass or fluffy-seed box for the native grass seed and the small-seed box for the forbs. However, if the seed has been cleaned and dechaffed it is common for seed companies to mix the seed and recommend it be planted together using the fluffy or grain box on a no-till drill. Fillers can be used when planting native grasses and forbs separately or when planting native grass and forb mixtures. Refer to the chart below for recommended fillers for the different seed boxes of a no-till drill.
Example:
We are planting a 10-acre field to a native warm-season grass and forb mixture using a no-till drill. The recommended seeding rate is 6 lbs/acre. The seed will be planted with the grain box of the drill, but the drill will only plant a minimum of 10 lbs/acre of our seed mixture.
10 acre field * 6 lbs/acre bulk seeding rate = 60 lbs of the seed mixture
Minimum seeding rate for the no-till drill is 10 lbs/acre = 10 lbs/acre * 10 acres = 100 lbs
We need to add a filler to increase the bulk weight of the seed mixture to be able to plant at the correct seeding rate. We added a 1:1 ratio (by weight) of cracked corn to our seed mixture:
60 lbs of seed + 60 lbs of cracked corn = 120 lbs of bulk weight for 10 acres
We now need to adjust our bulk seeding rate to account for the added crack corn.
120 lbs of bulk weight for 10 acres = 12 lbs/acre
We need to calibrate our drill to plant 12 lbs/acre in order to plant 6 lbs/ac of our initial seed mixture.
Generally, you should use a 1:1 ratio (by weight) of filler-to-seed, but in some cases you may need to use a higher ratio (e.g., 2:1, 3:1, or 4:1 filler-to-ratio) to achieve the correct seeding rate.
Using carriers when broadcasting native warm-season grass and forb mixtures
Broadcasting native warm-season grass and forbs mixtures is most commonly accomplished with a cyclone fertilizer spreader. These spreaders may have issues broadcasting the native grass and forb seed. The 2 main issues are: (1) the seed is not heavy enough to flow through the spreader and (2) the seeds of various size will settle and will not be spread evenly across the field. Carriers will add more bulk weight to the native grass seed and will help ensure the seed stays mixed across the field. Common carriers that are used with native grasses are cracked corn, pelletized lime, wheat, or oats. The recommended rates of common carriers are in the table below:
Table adapted from the publication Warm season grass establishment,
Indiana Department of Natural Resources, 2006.
Example:
We plan to broadcast a native grass and forb mixture on a 10-acre field. The recommended bulk seeding rate is 6 lbs/acre.
10 acre field * 6 lbs/acre bulk seeding rate = 60 lbs of the seed mixture
We need to add a carrier to the mixture to increase the bulk weight of the seed mixture. We plan to add 200 lbs of pelletized lime per acre:
200 lbs/ac of lime * 10 acres = 2000 lbs of lime
60 lbs of seed + 2000 lbs of pelletized lime = 2060 lbs of bulk weight for 10 acres
We now need to adjust our bulk seeding rate to account for the added pelletized lime.
2060 lbs of bulk weight for 10 acres = 206 lbs/acre
We need to calibrate the spreader to broadcast 206 lbs/acre in order to plant 6 lbs/acre of our seed mixture.
Conclusions
Planting native warm-season grass and forb mixtures at the correct rate is a critical step in ensuring a successful planting. Using fillers and carriers when establishing native warm-season grasses and forbs can help ensure the mixtures are planted at the proper rates, flow correctly through the seeding equipment, and ensure the seed is spread evenly across the field.
Other Resources:
Pure Live Seed: Calculations and Considerations for Wildlife Food Plots, Detailed Resource, Purdue Extension-FNR
Renovating Native Warm-Season Grass Stands for Wildlife: A Land Manager’s Guide, free pdf download
Calibrating a No-Till: FNR 556-WV, free pdf download
VIDEO: Calibrating a No-Till Drill for Conservation Plantings and Wildlife Food Plots
VIDEO: Frost Seeding to Establish Wildlife Food Plots & Native Grass and Forb Plantings
Printable Version:
PDF available for print: Seed Fillers and Carriers for Planting Native Warm-Season Grasses and Forbs (pdf 771.45) detail resource.
Moriah Boggess, Purdue Extension Wildlife Intern
Jarred Brooke, Purdue Extension Wildlife Specialist
Have you heard the old adage “proper planning prevents poor performance?” This adage applies perfectly to establishing food plots for wildlife; you just have to adjust the words, “Proper planting prevents poor food plot performance”.
When we talk about planting we include planting method, timing, depth, and planter calibration (see video), but we also include seeding rate. While there are many steps prior to planting to ensure a successful food plot, including taking a soil test, adjusting the soil fertility, and proper site preparation, determining the proper seeding rate based on pure live seed rather than bulk weight may be one aspect that people skip or ignore.
All agronomic seeds have a recommended seeding rate. This is the rate that maximizes forage or grain production and minimizes seed costs. Planting food plot seed too heavily is often a waste of money, because it increases your seed cost, but does not necessarily increase you forage or grain production. Planting food plots too lightly is an inefficient use of field space, opening areas up for weeds to overtake your planting, and could result in failure because of overbrowsing.
Seeding rate also varies by planting method and it’s important to make sure to follow the recommended seeding rate for the planting method you will be using. This rate is greater when broadcasting rather than when drilling or planting seed (broadcast rate: 75-100 lbs/ac vs. drilling rate: 50 lbs/ac for iron-clay cowpeas).
It is important to recognize the difference in how seed is sold (bulk weight) and how seeding rates are recommended (pure live seed [PLS]). When you buy seed from a supplier you are buying it in the form of bulk weight. Bulk weight is the total weight inside the bag, including the food plot seed, as well as the weight of the seed coating and other material (other crop seed and weed seed).
While seed is sold by bulk weight, seeding recommendations are commonly given as PLS rates. This distinction is important. For example, all the recommendations in A Guide to Wildlife Food Plots and Early Successional Plants by Dr. Craig Harper, University of Tennessee Extension are given in PLS rates. Pure live seed is the living seed of the intended crop that will germinate from a seed bag and accounts for the weight of the bag made up of impurities, weed seed, and other crop seed.
Percent PLS in a seed bag varies, depending on the amount of pure seed in the bag as well as the germination rate of that seed. It is important to know the difference between PLS and bulk weight so that you can calculate the amount of bulk weight you need to achieve the recommended PLS seeding rate.
Keep the distinction between bulk weight and PLS weight in mind when buying seed to ensure you purchase enough seed to cover the entire area of your food plot. In this article we will explain how to calculate PLS based on information given to you when you purchase seed.
Dissecting a seed tag – know what you’re getting before you purchase
Before we get too far into calculating PLS, it is important to understand what information you get when purchasing seed. Regardless of whether you buy a seed mix or a single species from a co-op, feed store, or sporting goods store, all agricultural seed sold in Indiana must come with a label – commonly called a seed tag. By law, that seed tag must contain these 12 items: 1) commonly accepted name (species) and variety, 2) lot number, 3) origin of seed, 4) company who labeled the seed, 5) percentage of pure seed (>5% of weight), 6) percentage of other crop seeds, 7) percentage of inert matter, 8) percentage of weed seed, 9) species and amount of any noxious weeds (if present), 10) germination rate, 11) hard seed, and 12) calendar month and year the bag was tested.
Important items to look for when purchasing seed are explained below.

As we can see from this red clover seed tag above, only 65% (32.5 lbs) of the 50 lb bag is actually red clover (Pure Seed). Most of the other weight is the inoculant (Coating Material; 34% or 17 lbs). Of the 32.5 lbs of red clover seed, 80% will germinate after planting (Germination). So, if you planted 15 lbs (bulk weight) from this bag (recommended PLS rate) you would actually be planting 9.75 lbs of pure seed, of which only 7.8 lbs would germinate. Meaning you would be planting a little over half the recommended rate. See why PLS is so important now?
Clover seed: bag half empty or half full?
It may seem odd to buy a 10 lb clover seed bag only to expect 5 lbs to actually grow, but in the case of legumes – like clovers – the 5 lbs in the bag that isn’t seed may actually save you money in the long run. Having as little as 50% PLS in a bag of clover is not uncommon, as much of the weight is made up of the seed coating, which is typically an inoculant. The inoculant on clover, soybeans, alfalfa, or other legumes is living bacteria that help fix nitrogen from the air and make it usable to plants. Planting inoculated seed – whether you inoculate it yourself or purchase pre-inoculated seed – reduces the amount of nitrogen fertilizer needed, ultimately saving you money on fertilizer application. The benefit of buying pre-inoculated seed is that it saves you the hassle of inoculating the seed yourself.
Calculating pure live seed (PLS)
Now that you know the importance of reading the seed tag before purchasing seed, you can now think about calculating PLS. All the information you need is right there on the tag. Using the example seed tag below locate the two items labeled “PURE SEED” and “GERMINATION”.
Our example is 98% pure seed, meaning in a 10 lb bag there are 9.8 pounds of pure iron-clay cowpea seed. The next number we will need is the germination rate; this will also be indicated as a percentage. Our example has an 85% germination rate.
To calculate percent PLS for this seed, multiply pure seed and germination and then multiply by 100 (example below). The result is the percentage of pure live seed for this bag of seed.
98% pure seed
85% germination
(0.98 x 0.85) x 100 = 83% (Percent PLS)
We now know that 83% of our bag is iron-clay cowpeas capable of germinating. Now that we have the PLS of our seed we can calculate the amount (lbs) of pure live seed in our bag. To do this, multiply the bulk seed weight by the percent PLS.
50 lb bag x 0.83 (Percent PLS) = 42 lbs PLS
For example, a 50 lb bag would equal 42 lbs of iron-clay cowpea seed that will germinate.
Determining bulk seed rate
Using the percent PLS information we can also find the amount of bulk seed necessary to plant at a recommended seeding rate. To do this, take the recommended seeding rate for your planting method and divide it by the percent PLS of the seed you are using (example below). The result is the bulk weight that you will need to plant per acre to achieve the recommended seeding rate.
50 lbs/acre = recommended seeding rate for iron-clay cowpeas when drilling
0.83 = percent PLS of seed being planted (from example above)
50/0.83 = 60 lbs/acre
In order to plant a cowpea food plot at the recommended rate of 50 lbs/acre, we will need to plant 60 lbs/acre from this particular seed bag.
Another benefit of calculating pure live seed? Comparing seed prices
Not all seeds are of the same quality. Some have lower pure seed and germination rates than others. For this reason, it is not always most cost effective to buy the cheapest bag of seed on the shelf. If you wish to compare pure live seed prices between seed bags, first divide the price of the bag by the weight of the bag. This will give you the price per pound of bulk seed (example below). Now divide this price per pound by the percent PLS of the seed. The result is the price per pound of PLS.
Seed Manufacturer X 10lbs
Pure seed 97%
Germination 92%
Price $33
PLS (0.97 x 0.92) x 100 = 89%
$/lb (bulk) 33/10 = $3.30/lb
$/lb (PLS) 3.3/0.89 = $3.70/lb
Seed Manufacturer Y 10lbs
Pure seed 91%
Germination 82%
Price $30
PLS (0.91 x 0.82) x 100 = 74.%
$/lb (bulk) 30/10 = $3/lb
$/lb (PLS) 3.0/0.74 = $4.05/lb
According to these calculations, seed manufacturer X is actually a better value than Y. Using this process, especially when purchasing seed in bulk, you can be a practical money saver.
Let’s Plant
Now that you know what all those numbers on the seed tag of your favorite food plot seed mean and you know how to calculate PLS, you will be able to plant your food plots at the most effective rate possible. Remember, regardless of whether you are planting acres of soybeans with a no-till drill or frost seeding a ½-acre clover patch, knowing how to calculate PLS will allow you to be more successful and save money when planting food plots.
Other Resources:
Seed Fillers and Carriers for Planting Native Warm-Season Grasses and Forbs, Detailed Resource, Purdue Extension-FNR
Renovating Native Warm-Season Grass Stands for Wildlife: A Land Manager’s Guide, free pdf download
VIDEO: Calibrating a No-Till Drill for Conservation Plantings and Wildlife Food Plots
VIDEO: Frost Seeding to Establish Wildlife Food Plots & Native Grass and Forb Plantings
Moriah Boggess, former Purdue Extension Wildlife Intern
Jarred Brooke, Purdue Extension Wildlife Specialist
Outbreaks of bovine tuberculosis have occurred sporadically around the world since the 1900s. In Bovine Tuberculosis in Wild White-tailed Deer, Bovine tuberculosis transmission, hosts, current status in Indiana, clinical signs, effects on deer populations, effect on white-tailed deer meat, management, and monitoring is extensively covered. If you’re a hunter or cattle producer, the information provided within this free publication would greatly benefit you!
Resources:
Bovine Tuberculosis in wild white-tailed deer, Got Nature?, Purdue Extension-FNR
Jarred Brooke, Extension Wildlife Specialist
Purdue Department of Forestry and Natural Resources
Teachers, parents and outdoor enthusiasts will want to download this free new Purdue Extension-Forestry and Natural Resources publication, The Great Clearcut Controversy. In this inquiry-based teaching unit, students use real scientific data to investigate how a bird community and individual forest animals respond to a clearcut timber harvest. In this investigation, students: use scientific inquiry to gain knowledge and answer questions; apply that knowledge to the engineering design process; and design a viable management solution given the constraints and tradeoffs they discover. All materials used in the three lessons are easily accessible and free.
Resources:
The Nature of Teaching, Purdue Extension
Got Nature? Podcast, Forestry and Natural Resources
Benefits of Connecting with Nature, The Education Store
Skye M Greenler, Graduate Research Assistant
Purdue University. Department of Forestry and Natural Resources
Mike Saunders, Associate Professor of Ecology and Natural Resources
Purdue University, Department of Forestry and Natural Resources
A recent study in Costa Rica by scientists at Princeton University-Department of Ecology and Evolutionary Biology and the University of Pennsylvania-Ecology & Biodiversity showed how agricultural waste could help successfully remediate barren tropical forest sites.
A deal between an orange juice manufacturer and advisors for the Área de Conservación Guanacaste was struck that allowed 1,000 truckloads (12 metric tons) of orange peels and pulp to be unloaded on nutrient-poor soils dominated by invasive grass within one of Costa Rica’s national parks in the mid-1990s. The land was left to rest for sixteen years. Now, the forest has begun the lengthy process of regeneration and some secondary forest regeneration has been observed. The peels were spread over 3 ha (7 acres) and scientists measured an increase of 176% in aboveground biomass. The distinction between the orange-fertilized land and the unfertilized areas was pronounced. Orange-fertilized areas demonstrated more tree biomass and species diversity and richer soils than their counterparts.

To enlarge photo click on image. Photo credits: Tim Treuer, Daniel Janzen, Winnie Hallwachs, & Leland Werden.
This research shows what could happen when the environmental community and industry find a medium where they can work together to come up with problem solutions. Perhaps, in future, similar uses can be found for agricultural excess here in the United States.
References:
Treuer TLH, Choi JJ, Janzen DH, Hallwachs W, Peréz-Aviles D, Dobson AP, Powers JS, Shanks LC, Werden LK, Wilcove DS. (2017) Low-cost agricultural waste accelerates tropical forest regeneration. Restoration Ecology, DOI: 10.1111/rec.12565
Resources:
Indiana Forestry & Woodland Owners Association
Purdue Extension-Forestry and Natural Resources workshops
Forest Improvement Handbook, The Education Store, Purdue Extension resource center
Shaneka Lawson, USDA Forest Service/HTIRC Research Plant Physiologist/Adjunct Assistant Professor
Purdue University Department of Forestry and Natural Resources
The calendar flipping over to October is a reminder the annual Autumn leaf color display is on its way. The perennial question is “how will the color be this year?” Predicting the quality of the fall display requires weighing several factors that may vary over time and across the landscape. In general, Indiana started the growing season wet and is ending it dry. The good growing conditions early this year have produced abundant leaf area on many trees. As we have dried out late in the season, some trees are experiencing stress that may cause leaves to turn color and drop early, or to simply turn brown. Drought may also delay the color change by a week or more in some cases. Leaves that were attacked by insects or disease may also drop early or provide very little color. I noticed Japanese beetles seemed to be more active than normal this year, skeletonizing leaves on preferred plants like linden and Virginia creeper. Local weather patterns can also influence color intensity in a positive way. Sunny days and cool nights late in the summer and early fall can enhance the production of anthocyanin – a pigment produced in some trees that provides bright red, maroon and purple tones to the fall color palate.
This brings us to another variable: different species of trees may produce different colors, timing, and duration of fall color. Some species like sassafras, sumac, black and sweet gum, and sugar and red maple are famous for bright fall color. Some species like elms, buckeye and walnut may simply turn brown or drop early with little color display. Different individual trees may also vary due to genetic differences, growing conditions and tree health. For example, some sugar maple located in open areas or on the edge of a woodlot, receiving lots of sunlight, may regularly produce vibrant oranges and reds, while nearby sugar maple in the shade of the forest will turn a subdued yellow, lacking the sugar reserves produced by their neighbors in the sun.
Leaf color change is also the result of a very predictable process based on the longer night period as summer slips into fall. The production of green chlorophyll pigments slows and finally stops as the nights become longer and cooler, exposing the yellows and oranges of carotenoids and reds and maroons of anthocyanins. This process also starts forming a zone of separation between the leaf and branch that ultimately brings the leaves to the ground, often with the help of wind and rain.
My best answer to those asking for a fall color prediction is another set of questions: how was your weather, what species of trees do you have, and how much sunlight do they receive?
Resources:
Why Leaves Change Color, The Education Store, Purdue Extension
Why Leaves Change Color, USDA Forest Service, Northeastern Area
Fifty Common Trees of Indiana
An Introduction to Trees of Indiana
Native Trees of the Midwest, The Education Store
Shrubs of Indiana CD: Their Identification and Uses, The Education Store
Shrubs and Woody Vines of Indiana and the Midwest, The Education Store
Trees of Indiana CD, The Education Store
Lenny Farlee, Sustaining Hardwood Extension Specialist
Purdue Department of Forestry and Natural Resources
Spring and Fall is prime time for improving your property with new trees. They provide many benefits which everyone can share. Trees mean more attractive landscapes, lower energy bills and a healthier environment. However, just planting a tree without some thought and planning can create a liability rather than an asset to your site. Wise planning is essential to ensure the new trees meet your design needs and functional solutions as well. Follow these basic tips to get your tree started right and make it a long-lasting sustainable planting. For more information, download the free publication Tree Installation: Process and Practice.
Right Tree-Right Place.
Location, location, location! Planning before planting can help ensure that the right tree is planted in the right place. Proper tree selection and placement enhances your property value, prevents costly and sometimes unsightly maintenance with trimming, and lowers the risk of damage to your home and property. In some instances, trees are the innocent victim of poor planting locations and must be removed. Always allow room for growth! Also, consider native trees or those trees with fewer pests which can attack your tree. Large trees include Kentucky Coffeetree, Bur Oak and Hardy Rubber Tree. Medium-sized trees can include Japanese Pagoda Tree, Sourwood, Katsura Tree and Golden Raintree. Finally, for areas with less room, consider Serviceberry, Ironwood, Amur Maackia or Hop Tree. These are just a few of the many trees which can be chosen for your situation.
Look Up, Look Down, and Look All Around!
Regardless if the planting is in the front yard or the back yard of the home or business, be sure there will be no interference with utilities; Call 811 before you dig. It will prevent costly mistakes and maybe a life. In addition, if the tree is going to be planted along the street, typically, there is an ordinance requiring a permit to plant in the right of way. This helps Urban Forestry administration keep up the street tree inventory and allows the ISA Certified Arborists on staff a chance to offer free advice to help in the planting decisions.
It Comes from Good Stock…
Choose the tree twice, meaning get the right species for your location; then, make an informed choice on the nursery stock. Be sure the function of the tree is understood and choose the right tree for the location. Shade? Flowers? Screening? Sound Barrier? Trees can be used as tools to work for you on the site. “You get what you pay for” applies to nursery stock as well. Purchase plant material from a reputable source and get a professional opinion on the tree species for your application. One hint, if it is a fast growing tree, it probably won’t last long. See our video for tree selection tips.
This Hole is a Home!
It is a permanent home for the trees… understand the planting site prior to planting. Determine soil type and pH, drainage and exposure to the sun. If the tree isn’t naturally suited to the planting place, it doesn’t have a chance. Planting depth is a major tree planting concern. Be sure to find the “root flare” when establishing the final grade of the tree. Drainage is crucial to survival. Use the two-hour test. Dig the hole, fill it with water. If the hole is empty upon returning, there is suitable drainage for any tree. Plant the tree properly and at the proper depth, you only get one chance… Don’t dig a $10 hole for a $100 tree. See our video on tree planting tips.
Keep Good Care of the Investment.
Once the tree is in the ground, take good care of it. At least an inch of water per week to keep it growing vigorously, apply clean, hardwood mulch on the root zone to keep soils cool and moist, but never exceed three inches in depth. Remember to remove any tags on the tree and don’t forget to remove the twine from around the trunk. Don’t worry about the fertilizer at planting time, wait until next year, after the tree has gotten settled in to its new home. Enjoy your new addition to the home and landscape!
Resources:
Tree Selection for the “Un-natural” Environment, The Education Store, Purdue Extension
Tree Support Systems, The Education Store, Purdue Extension
Tree Installation: Process and Practices, The Education Store, Purdue Extension
Planting Your Tree Part 1: Choosing Your Tree, video, The Education Store, Purdue Extension
Lindsey Purcell, Urban Forestry Specialist
Purdue University, Department of Forestry and Natural Resources
Sericea lespedeza is arguably one of the most problematic invasive species of old fields, prairies and other early successional areas managed for wildlife. Sericea lespedeza is a perennial legume native to eastern Asia that was originally promoted for erosion control, cattle forage, cover and food for wildlife. But as with many plant introductions during the early and mid-19th century (e.g., multiflora rose, autumn olive, and bush honeysuckle), the original beliefs – while well intentioned – were ill fated and short-sided. Sericea has become invasive, is considered noxious in many states, and can be found from Massachusetts to Nebraska and from Florida to Ontario.
Here are 3 problems with sericea and 3 tools for control.
Problems:
If you find sericea in fields that you manage, working quickly to stop seed production and kill the existing plants will be the most effective way to control an invasion.
Web Resources:
Herbicides to control sericea lespedeza, Southeastern Association of Fish & Wildlife Agencies
Effects of Growing-Season Prescribed Burning on Vigor of Sericea Lespedeza in the Kansas Flint Hills: I. Suppression of Seed Production and Canopy Dominance, Kansas Agricultural Experiment Station Research Reports
Other Resources:
If Your Native Grasses Look Like This, It’s Time for Management, Got Nature?, Purdue Extension-FNR
Invasive plants: impact on environment and people, The Education Store, Purdue Extension
Jarred Brooke, Extension Wildlife Specialist
Purdue University Department of Forestry and Natural Resources
Due to reasons such as pruning, location of growth and species characteristics, trees can grow in ways that don’t endorse long-term health and safety. To counter trees growing in unsafe ways, cabling, bracing, guying, or props can be utilized to prevent branch or whole-tree failure. These tree support systems reinforce critical areas of the tree by limiting the movement of branches or leaders. In the publication titled Large Tree Cabling and Bracing, FNR-550-W, common structural deficiencies in trees and the tree support devices used to prevent problems caused by those deficiencies are described and covered.
Resources:
Tree Support Systems, The Education Store, Purdue Extension
Tree Planting Part 1: Choosing a Tree, The Education Store, Purdue Extension
Tree Pruning: What Do Trees Think?, The Education Store, Purdue Extension
Forest Improvement Handbook, The Education Store, Purdue Extension
Got Nature?, Purdue Extension, Forestry and Natural Resources
Lindsey Purcell, Urban Forestry Specialist
Purdue University, Department of Forestry and Natural Resources